Camponotus herculeanus (Linnaeus, 1758) valid
Formica herculeana Linnaeus, 1758: 579 (q.) EUROPE. Palearctic.
Type notes: Type-locality Sweden, after Linnaeus, 1761: 426.
  • [Misspelled as herculanea by Latreille, 1809: 126, Smith, 1858a: 10; misspelled as hepculeanus by Kuznetsov-Ugamsky, 1929a: 17; misspelled as helcureanus by Teranishi, 1940: 71, and subsequent pages.].
  • Lepeletier de Saint-Fargeau, 1835: 209 (w.m.); Wheeler & Wheeler, 1953e: 185 (l.).
  • Combination in Camponotus: Mayr, 1861: 36 (in key).
  • Combination in Camponotus (Camponotus): Forel, 1914a: 259.
  • Status as species: Linnaeus, 1761: 426; Scopoli, 1763: 312; Linnaeus, 1767: 962; Fabricius, 1775: 391; Fabricius, 1782: 488; Fabricius, 1787: 307; Gmelin, 1790: 2797; Christ, 1791: 514; Olivier, 1792: 491; Fabricius, 1793: 349; Latreille, 1798: 33; Fabricius, 1804: 395; Jurine, 1807: 272; Gravenhorst, 1807: 286; Latreille, 1809: 126; Leach, 1815: 147; Latreille, 1817a: 96; Billberg, 1820: 104; Brullé, 1833: 326; Losana, 1834: 309; Lepeletier de Saint-Fargeau, 1835: 209; Zetterstedt, 1838: 448; Schilling, 1839: 52; Nylander, 1846a: 894; Nylander, 1846b: 1044; Foerster, 1850a: 9; Schenck, 1852: 123; Mayr, 1855: 308 (redescription); Nylander, 1856b: 56; Gredler, 1858: 3; Smith, 1858a: 10, 53; Roger, 1859: 229; Duméril, 1860: 929; Mayr, 1861: 36 (in key); Meinert, 1861: 309; Mayr, 1863a: 399; Roger, 1863b: 1; Cresson, 1865b: 426; Dours, 1873: 164; Forel, 1874: 39 (in key); André, 1874b: 176 (in key); Emery, 1878: 44; Emery & Forel, 1879: 447; Forel, 1879a: 56; Provancher, 1881b: 354; André, 1882b: 142 (in key); Provancher, 1883: 597; Mayr, 1886d: 419; Cresson, 1887: 255; Provancher, 1887: 228 (in key); Nasonov, 1889: 9; Forel, 1892j: 306; Dalla Torre, 1893: 233; Emery, 1893k: 674; Ruzsky, 1896: 67; Emery, 1896j: 372 (in list); Ruzsky, 1902d: 5; Forel, 1902k: 699; Ruzsky, 1903c: 205; Forel, 1904c: 381; Ruzsky, 1905b: 214; Wasmann, 1906: 111; Forel, 1907h: 19; Emery, 1908a: 182; Wheeler, 1908i: 625; Bondroit, 1910: 487; Santschi, 1911d: 7; Karavaiev, 1912b: 592; Forel, 1914a: 266; Stitz, 1914: 95; Bruch, 1914: 228; Ruzsky, 1914b: 100; Bruch, 1915: 535; Forel, 1915d: 68 (in key); Ruzsky, 1915b: 4; Donisthorpe, 1915f: 347; Emery, 1916a: 225; Escherich, 1917: 330 (in key); Bondroit, 1918: 70; Nadig, 1918: 339; Menozzi, 1922d: 142; Soudek, 1922b: 95; Kuznetsov-Ugamsky, 1923b: 241; Müller, 1923b: 159; Wheeler, 1923c: 5; Vashkevich, 1924b: 146; Emery, 1925d: 72; Kiseleva, 1925: 73; Ruzsky, 1925a: 287; Essig, 1926: 868; Karavaiev, 1926e: 191; Ruzsky, 1926: 108; Stärcke, 1926a: 119 (in key); Karavaiev, 1927a: 295; Donisthorpe, 1927c: 400; Menozzi, 1927b: 92; Kuznetsov-Ugamsky, 1929b: 36; Karavaiev, 1930b: 147; Arnol'di, 1933a: 602 (in key); Stitz, 1934: 4; Grandi, 1935: 102; Karavaiev, 1936: 177 (redescription); Ruzsky, 1936: 89; Kôno & Sugihara, 1939: 10; Teranishi, 1940: 71; Novák & Sadil, 1941: 110 (in key); Holgersen, 1942b: 10; Holgersen, 1943c: 172 (in key); Holgersen, 1944a: 179; Ruzsky, 1946: 69; Creighton, 1950a: 366; Donisthorpe, 1950e: 1066; Röszler, 1950: 210; Yasumatsu & Brown, 1951: 30; Chapman & Capco, 1951: 220; Consani & Zangheri, 1952: 43; Cole, 1954f: 271; Ceballos, 1956: 312; Smith, 1958c: 142; Pisarski, 1961: 153; Dlussky, 1962: 181; Collingwood, 1962c: 220; Bernard, 1967a: 340 (redescription); Arnol'di, 1967: 1819 (redescription); Smith, 1967a: 366; Kutter, 1968b: 60; Collingwood & Yarrow, 1969: 81; Dlussky & Pisarski, 1970: 86; Baroni Urbani, 1971c: 175; Collingwood, 1971: 163; Banert & Pisarski, 1972: 352; Tarbinsky, 1976: 148 (redescription); Aktaç, 1977: 125; Van Boven, 1977: 131; Francoeur, 1977b: 207; Kutter, 1977c: 204; Yensen et al., 1977: 183; Collingwood, 1978: 91 (in key); Collingwood, 1979: 90; Smith, 1979: 1426; Allred, 1982: 454; Wheeler & Wheeler, 1986g: 60; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 283 (in key); Nilsson & Douwes, 1987: 68; MacKay et al., 1988: 105; Wang et al., 1989a: 224 (in key); Morisita et al., 1991: 41; Wang, 1992: 681; Atanassov & Dlussky, 1992: 210; Arakelian, 1994: 85; Radchenko, 1994b: 116 (in key); Wang & Wu, 1994: 31 (in key); Wheeler et al., 1994: 305; Bolton, 1995b: 103; Douwes, 1995: 92; Poldi et al., 1995: 7; Wu & Wang, 1995a: 182; Radchenko, 1996e: 1202 (in key); Espadaler, 1997g: 27; Radchenko, 1997b: 555; Gallé et al., 1998: 216; Czechowski et al., 2002: 95; Mackay & Mackay, 2002: 290; Zhang & Zheng, 2002: 218; Coovert, 2005: 166; Hansen & Klotz, 2005: 82; Radchenko, 2005b: 158; Csősz & Markó, 2005: 228; Karaman & Karaman, 2005: 58; Bračko, 2006: 145; Markó et al., 2006: 66; Petrov, 2006: 109 (in key); Schultz et al., 2006: 203; Bračko, 2007: 19; Seifert, 2007: 262; Werner & Wiezik, 2007: 143; Zryanin & Zryanina, 2007: 232; Gratiashvili & Barjadze, 2008: 131; Casevitz-Weulersse & Galkowski, 2009: 479; Lapeva-Gjonova et al., 2010: 43; Boer, 2010: 18; Csősz et al., 2011: 58; Karaman, 2011b: 70; Legakis, 2011: 30; Ran & Zhou, 2011: 67; Borowiec & Salata, 2012: 476; Czechowski et al., 2012: 240; Ellison et al., 2012: 121; Guénard & Dunn, 2012: 28; Kiran & Karaman, 2012: 7; Karaman & Aktaç, 2013: 52 (in key); Borowiec, 2014: 32; Lebas et al., 2016: 126; Radchenko, 2016: 328; Salata & Borowiec, 2018c: 43; Schär et al., 2018: 6; Seifert, 2018: 256; Mackay, 2019: 213 (redescription).
  • [Note: Seifert, 2019a: 1, reports Camponotus herculeanus × Camponotus ligniperdus worker hybrids.].
  • Senior synonym of Camponotus herculeanus altaica: Schär et al., 2018: 6.
  • Senior synonym of Camponotus atrus: Nylander, 1846a: 894; Foerster, 1850a: 9; Nylander, 1856b: 56; Smith, 1858a: 10; Mayr, 1863a: 399; Roger, 1863b: 1; Forel, 1874: 96 (in list); Emery & Forel, 1879: 447; Dalla Torre, 1893: 234; Emery, 1925d: 72; Karavaiev, 1936: 178; Bolton, 1995b: 103; Radchenko, 2016: 328.
  • Senior synonym of Camponotus herculeanus caucasicus: Arakelian, 1994: 85; Bolton, 1995b: 103; Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus herculeanus eudokiae: Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus intermedia: Nylander, 1846a: 894; Foerster, 1850a: 9; Nylander, 1856b: 56; Smith, 1858a: 10; Mayr, 1863a: 399; Roger, 1863b: 1; Emery & Forel, 1879: 447; Emery, 1925d: 72; Karavaiev, 1936: 178; Bolton, 1995b: 103; Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus herculeanus jacuticus: Yasumatsu & Brown, 1951: 35; Schär et al., 2018: 6.
  • Senior synonym of Camponotus herculeanus montanus: Emery, 1925d: 72; Karavaiev, 1936: 178; Yasumatsu & Brown, 1951: 30; Arnol'di, 1967: 1819; Arakelian, 1994: 85; Bolton, 1995b: 103; Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus herculeanus nadigi: Yasumatsu & Brown, 1951: 30; Baroni Urbani, 1971c: 176; Bolton, 1995b: 103; Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus herculeanus sachalinensis: Yasumatsu & Brown, 1957: 49; Schär et al., 2018: 6.
  • Senior synonym of Camponotus herculeanus shitkowi: Emery, 1925d: 72; Karavaiev, 1936: 178; Arnol'di, 1967: 1819; Bolton, 1995b: 103; Radchenko, 1997b: 555; Radchenko, 2016: 328.
  • Senior synonym of Camponotus whymperi: Creighton, 1950a: 367; Yasumatsu & Brown, 1951: 35; Smith, 1958c: 142; Smith, 1979: 1426; Bolton, 1995b: 103; Mackay, 2019: 214.
  • Material of the unavailable name Camponotus herculeanus shitkowi intermedius referred here by Radchenko, 1997b: 555.
Junior synonyms
Obsolete combinations
Citations