Camponotus - AntCat

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Camponotus Mayr, 1861 valid masculine

Camponotus Mayr, 1861: 35 Type-species: Formica ligniperda (obsolete combination of Camponotus ligniperda), by subsequent designation of Bingham, 1903: 347.

[Type-species not Formica herculeana, unjustified subsequent designation by Forel, 1914a: 259; repeated by Arnold, 1922: 612.]
Camponotus in Formicinae: Mayr, 1861: 35 [Formicidae]; Mayr, 1862: 652 (in key) [Formicidae]; Mayr, 1865: 6 [Formicidae]; Mayr, 1868b: 26 [Formicidae]; Dours, 1873: 164 [Formicidae]; Forel, 1874: 37 [Formicidae].
Camponotus in Camponotinae: Forel, 1878c: 367 [Camponotidae]; Emery & Forel, 1879: 447 [Camponotidae]; Forel, 1892k: 223 [Camponotidae]; Dalla Torre, 1893: 219 [Camponotinae]; Emery, 1896e: 188 [Camponotinae].
Camponotus in Camponotinae, Camponotini: Forel, 1886h: 141; Forel, 1893b: 165; Forel, 1895b: 101; Emery, 1895l: 772; Forel, 1899b: 130; Ashmead, 1905c: 384; Wheeler, 1910a: 144; Forel, 1912j: 89; Wheeler, 1915i: 138; Emery, 1916a: 222; Forel, 1917: 250; Arnold, 1922: 610.
Camponotus in Formicinae, Formica genus group: Agosti, 1991: 295.
Camponotus in Formicinae, Camponotini: Bondroit, 1918: 65; Wheeler, 1922: 703; Emery, 1925d: 59; subsequent authors; Hölldobler & Wilson, 1990: 18; Brandão, 1991: 332; Bolton, 1995b: 23; Bolton, 2003: 27, 112; Ward et al., 2016: 345.
Camponotus as subgenus of Formica: Smith, 1871b: 306.
Camponotus as genus: all authors except Smith, 1871a: 306.
Camponotus as senior synonym of †Paleosminthurus: Snelling, R.R. (pers. comm. to B. Bolton, 2004). [Synonymy remained unpublished at his death in 2008, but is accepted provisionally here.]
Camponotus as senior synonym of Dolophra: Bolton, 1995b: 23; Ward & Boudinot, 2021: 44.
[Note (i): all subgenera were given as provisional junior synonyms of Camponotus by Brown, 1973b: 179-185. The list was repeated by Hölldobler & Wilson, 1990: 18, and Tang et al., 1995: 112, with all subgenera listed as junior synonyms. They reverted to subgeneric status in Brandão, 1991: 330 (Neotropical), Bolton, 1994: 50 (world), and Bolton, 2003: 113 (world). The entry of Myrmophyma and Thlipsepinotus under the synonymy of Camponotus by Taylor & Brown, 1985: 109, is not accepted as confirmation as not all taxa were included.]
[Note (ii): the subgeneric classification is retained here pending formal revision. While some subgenera appear monophyletic it is clear that several are patently artificial, including the largest subgenus, Camponotus (Tanaemyrmex).]

Dolophra Wu & Wang, 1994
†Paleosminthurus Pierce & Gibron, 1962

Mayr, 1861: 35 (Europe species key); Mayr, 1865: 6, 7 (Camponotus, Colobopsis diagnosis); Mayr, 1867a: 67 (Colobopsis diagnosis); Mayr, 1868b: 27 (Baltic Amber species key); Mayr, 1870a: 373 (Colombia + Panama (= New Grenada) species key); Mayr, 1870b: 941 (Colobopsis species key); André, 1874b: 175 (Europe species key); Forel, 1874: 38 (Switzerland species key); Mayr, 1876: 58 (Australia species key); Mayr, 1878: 869 (all Dendromyrmex species key (as part of Camponotus)); Forel, 1878c: 367, 368 (Camponotus, Colobopsis diagnoses); André, 1882b: 138 (Europe, Algeria species key); André, 1882c: 159 (Europe, Algeria Colobopsis species key); Provancher, 1887: 228 (Canada species key); Nasonov, 1889: 57 (Russia species key); Forel, 1891c: 71, 216 (Madagascar species key); Forel, 1892k: 223 (India, Sri Lanka species key); Emery, 1893k: 667 (North America species key); Forel, 1893c: 435 (India, Sri Lanka Camponotus (Colobopsis) species key); Emery, 1895l: 772 (synoptic classification); Emery, 1896j: 370 (species groups); Emery, 1903: 65 (Camponotus novogranadensis group key); Bingham, 1903: 343, 348 (India, Sri Lanka, Burma Colobopsis, Camponotus species keys); Ruzsky, 1905b: 185 (Russian Empire species key); Emery, 1908a: 183 (Palaearctic Camponotus herculeanus, Camponotus maculatus groups species key); Bondroit, 1910: 486 (Belgium species key); Wheeler, 1910g: 297 (North America species key); Forel, 1914a: 259, 266 (subgenera diagnoses); Stitz, 1914: 94 (Central Europe species key); Santschi, 1915c: 266 (Camponotus (Myrmotrema) species key); Emery, 1916a: 228 (Italy species key); Wheeler, 1916r: 600 (U.S.A., Connecticut species key); Mann, 1916: 489 (Dendromyrmex, all species key); Bondroit, 1918: 67 (France, Belgium species key); Emery, 1920b: 235 (subgenera diagnoses); Emery, 1920d: 8 (Old World Camponotus maculatus group, checklist, key); Wheeler, 1921a: 16 (subgenera, notes); Santschi, 1921f: 310 (subgenera, notes); Mann, 1921: 489, 499 (Fiji Is Camponotus (Myrmogonia), Camponotus (Colobopsis) species keys); Wheeler, 1922: 232 (Camponotus diagnosis); Wheeler, 1922: 703, 704 (short history of subgeneric classification, subgenera synopsis); Wheeler, 1922: 704 (Camponotus (Camponotus)); Wheeler, 1922: 992 (Phasmomyrmex diagnosis); Arnold, 1922: 610, 612, 619, 662, 673 (diagnosis, South Africa subgenera synopsis, South Africa Camponotus (Tanaemyrmex), Camponotus (Myrmophyma), Camponotus (Myrmopsamma) species keys); Arnold, 1924: 683, 700, 709, 724, 735 (South Africa Camponotus (Myrmosericus), Camponotus (Myrmamblys), Camponotus (Orthonotomyrmex), Camponotus (Myrmotrema), Camponotus (Myrmepomis) species keys); Emery, 1925e: 62 (Palaearctic Camponotus lateralis group, key); Emery, 1925d: 57, 58 (Phasmomyrmex, Phasmomyrmex (Myrmorhachis), Phasmomyrmex (Myrmacantha) diagnoses); Emery, 1925d: 59 (Camponotus diagnosis, subgenera key); Emery, 1925d: 69 (Camponotus (Dinomyrmex) diagnosis); Emery, 1925d: 70 (Camponotus (Myrmoxygenys), Camponotus (Myrmopsamma) diagnoses); Emery, 1925d: 71 (Camponotus (Camponotus) diagnosis); Emery, 1925d: 75 (Camponotus (Tanaemyrmex) diagnosis); Emery, 1925d: 104 (Camponotus (Myrmosericus) diagnosis); Emery, 1925d: 107 (Camponotus (Myrmothrix) diagnosis); Emery, 1925d: 109 (Camponotus (Myrmophyma) diagnosis); Emery, 1925d: 112 (Camponotus (Myrmosaulus) diagnosis); Emery, 1925d: 114 (Camponotus (Myrmopytia) diagnosis); Emery, 1925d: 115 (Camponotus (Karavaievia) diagnosis); Emery, 1925d: 116 (Camponotus (Myrmentoma) diagnosis); Emery, 1925d: 121 (Camponotus (Mayria) diagnosis); Emery, 1925d: 123 (Camponotus (Myrmonesites) diagnosis); Emery, 1925d: 124 (Camponotus (Orthonotomyrmex) diagnosis); Emery, 1925d: 126 (Camponotus (Myrmepinotus) diagnosis); Emery, 1925d: 127 (Camponotus (Myrmopiromis) diagnosis); Emery, 1925d: 130 (Camponotus (Myrmotrema) diagnosis); Emery, 1925d: 133 (Camponotus (Myrmisolepis) diagnosis); Emery, 1925d: 134 (Camponotus (Myrmotarsus) diagnosis); Emery, 1925d: 135 (Camponotus (Myrmoplatys) diagnosis); Emery, 1925d: 136 (Camponotus (Myrmamblys) diagnosis); Emery, 1925d: 142 (Camponotus (Rhinomyrmex) diagnosis); Emery, 1925d: 143 (Camponotus (Myrmogonia) diagnosis); Emery, 1925d: 144 (Camponotus (Colobopsis) diagnosis); Emery, 1925d: 151 (Camponotus (Myrmosphincta) diagnosis); Emery, 1925d: 152 (Camponotus (Myrmaphaenus) diagnosis); Emery, 1925d: 157 (Camponotus (Pseudocolobopsis) diagnosis); Emery, 1925d: 159 (Camponotus (Manniella) diagnosis); Emery, 1925d: 160 (Camponotus (Hypercolobopsis) diagnosis); Emery, 1925d: 161 (Camponotus (Myrmostenus), Camponotus (Myrmobrachys) diagnoses); Emery, 1925d: 166 (Camponotus (Myrmocladoecus) diagnosis); Emery, 1925d: 167 (Camponotus (Myrmeurynota) diagnosis); Emery, 1925d: 168 (Camponotus (Myrmodirhachis) diagnosis); Emery, 1925d: 169 (Camponotus (Myrmomalis), Camponotus (Myrmoplatypus) diagnoses); Emery, 1925d: 170 (Camponotus (Myrmepomis) diagnosis); Emery, 1925d: 172 (Dendromyrmex diagnosis); Santschi, 1926a: 16, 21 (Camponotus (Myrmopelta) diagnosis, species key); Karavaiev, 1927d: 275 (Ukraine species key); Arnol'di, 1933a: 601 (Russia species key); Menozzi, 1933b: 81 (Israel species key); Menozzi, 1935c: 330 (Chile species key); Karavaiev, 1936: 175 (Ukraine species key); Finzi, 1936: 190 (Egypt species key); Menozzi, 1939a: 316 (Himalaya, Tibet species key); Santschi, 1939d: 80 (North Africa Camponotus compressus group key); Stitz, 1939: 236 (Germany species key); Kratochvíl, 1941b: 109 (Central Europe species key); Novák & Sadil, 1941: 109 (Central Europe species key); Cole, 1942: 387 (U.S.A., Utah species key); Donisthorpe, 1942h: 248 (Camponotus (Orthonotomyrmex) notes); Holgersen, 1943c: 174 (Norway species key); Holgersen, 1944a: 199 (Norway species key); Buren, 1944a: 293 (U.S.A., Iowa species key); Creighton, 1950a: 362 (Nearctic species keys); Yasumatsu & Brown, 1951: 29 (Camponotus herculeanus complex); Kusnezov, 1952f: 192 (Argentina species key); Yasumatsu & Brown, 1957: 45 (Camponotus herculeanus complex); Pisarski, 1961: 175 (Poland species key); Gregg, 1963: 655 (U.S.A., Colorado species key); Wheeler & Wheeler, 1963: 163 (U.S.A., North Dakota species key); Arnol'di, 1967: 1817 (former U.S.S.R. Camponotus (Camponotus) species key); Bernard, 1967a: 330 (diagnosis, Western Europe species key); Hashmi, 1973b: 1 (Camponotus (Myrmothrix) all species revision, key); Alayo, 1974: 27 (Cuba species key); Snelling & Hunt, 1976: 117 (Chile species key); Tarbinsky, 1976: 144 (Kyrghyzstan species key); Van Boven, 1977: 130 (Belgium species key); Kutter, 1977c: 191 (Switzerland species key); Arnol'di & Dlussky, 1978: 551 (former European U.S.S.R. species key); Collingwood, 1978: 90 (Iberian Peninsula species key); Collingwood, 1979: 87 (Fennoscandia, Denmark species key); Allred, 1982: 445 (U.S.A., Utah species key); Gösswald, 1985: 264 (Germany species key); Collingwood, 1985: 275 (Saudi Arabia species key); Dumpert, 1986: 571 (Camponotus (Karavaievia) species key); Wheeler & Wheeler, 1986g: 60 (U.S.A., Nevada species key); Agosti & Collingwood, 1987b: 282 (Balkans species key); Snelling, 1988: 57 (Camponotus (Myrmentoma) Nearctic species key); Wang et al., 1989a: 222 (China species key); Dlussky et al., 1990: 125 (Turkmenistan species key); Kupyanskaya, 1990a: 164 (Far Eastern Russia species key); Morisita et al., 1991: 38 (Japan species key); Terayama, 1991b: 165 (Japan Camponotus (Paramyrmamblys) species); Atanassov & Dlussky, 1992: 207 (Bulgaria species key); Wang & Wu, 1994: 28 (China species key); Radchenko, 1994b: 116 (South Siberia species key); Arakelian, 1994: 82 (Armenia species key); Bolton, 1995a: 1048 (census); Douwes, 1995: 92 (Sweden species key); Dumpert et al., 1995: 104 (Camponotus (Karavaievia) species key); Kupyanskaya, 1995: 352 (Far Eastern Russia species key); Wu & Wang, 1995a: 168 (China species key); Collingwood & Agosti, 1996: 370 (Saudi Arabia species key); Seifert, 1996b: 170 (Central Europe species key); Radchenko, 1996e: 1195 (Asiatic Palaearctic species key); Cagniant, 1996b: 87 (Morocco species key); McArthur & Adams, 1996: 18 (Australia Camponotus nigriceps group, key); Dlussky, 1997a: 55 (in Baltic amber genera key); MacKay, 1997a: 194 (Camponotus (Myrmostenus) species key); Mackay & MacKay, 1997b: 319 (Camponotus montivagus complex, key); Robertson & Zachariades, 1997: 2 (Camponotus fulvopilosus group, key); Snelling & Torres, 1998: 9 (Puerto Rico, Virgin Is species key); Collingwood & Prince, 1998: 24 (Portugal species key); Terayama, 1999b: 27 (Japan species key); Shattuck, 1999: 91 (Australia synopsis); Snelling, 2000: 607 (Camponotus montivagus complex, key); Zhou, 2001a: 200 (China, Guangxi species key); McArthur & Shattuck, 2001: 27 (Australia Camponotus macrocephalus group, key); Czechowski et al., 2002: 153 (Poland species key); Fernández, 2002a: 58 (Camponotus (Dendromyrmex) diagnosis, all species revision, key); Mackay & Mackay, 2002: 248 (U.S.A., New Mexico species key); Shattuck & McArthur, 2002: 65 (Australia Camponotus wiederkehri group key); Bolton, 2003: 112 (Camponotus synopsis); Bolton, 2003: 122 (Phasmomyrmex synopsis); Shattuck, 2005: 3 (Camponotus aureopilus group key); Radchenko, 2005b: 188 (North Korea species key); Hansen & Klotz, 2005: 75 (North American “Carpenter Ants” key); Snelling, 2006: 96 (U.S.A. Camponotus festinatus complex key); Karmaly & Narendran, 2006: 29 (India species key); Dumpert et al., 2006: 77 (Camponotus (Karavaievia) species key); Andersen, 2007: 32 (Australia species groups); MacGown et al., 2007: 9 (southeast U.S.A. species key); McArthur, 2007b: 290 (Australia species key); Seifert, 2007: 152 (North and Central Europe species key); Zettel & Zimmermann, 2007: 19 (Forelophilus diagnosis, all species revision, key); Aibek & Yamane, 2009: 98 (Mongolia species key); Shattuck & Janda, 2009: 252 (Camponotus aureopilus group key); Mackay & Delsinne, 2009: 496 (New World Camponotus maculatus complex key); Terayama, 2009: 216 (Taiwan species key); Heterick, 2009: 60 (south-western Australia species key); Ionescu-Hirsch, 2010: 59 (Israel species key); McArthur, 2010: 18 (south Australia species key); Karaman et al., 2011: 191 (Turkey, Kaz Mountains species key); McArthur, 2012: 9 (Camponotus (Colobopsis) species review); Czechowski et al., 2012: 380 (Poland species key); Sarnat & Economo, 2012: 44 (Fiji Is species key); Karaman & Aktaç, 2013: 49 (Turkey species key); Dlussky & Perfilieva, 2014: 419 (rock fossil diagnosis); Bharti & Wachkoo, 2014a: 7 (Shivalik hills of Indian Himalaya species key); Dlussky & Putyatina, 2014: 249 (diagnosis, compression fossils); McArthur, 2014: 24 (Australia species key); Baccaro et al., 2015: 178 (genus in Brazil); Fisher & Bolton, 2016: 224 (Camponotus diagnosis); Fisher & Bolton, 2016: 362 (Phasmomyrmex diagnosis); Radchenko, 2016: 325 (Ukraine species key); Rakotonirina et al., 2016: 99 (Malagasy Camponotus edmondi group key); Ward et al., 2016: 341 (Camponotus/Colobopsis separation key); Clouse et al., 2016: 139 (Micronesia species); Rakotonirina et al., 2017: 215 (Malagasy Camponotus grandidieri and Camponotus niveosetosus groups key); Rasoamanana et al., 2017: 130 (Camponotus (Myrmopytia) species key); Cantone, 2017: 151 (brief male diagnosis); Rakotonirina & Fisher, 2018: 14 (Malagasy Camponotus (Mayria) species revision, key); Zettel et al., 2018: 129 (Camponotus inquilinus group key); Seifert, 2019b: 13 (Camponotus lateralis group, Europe, Asia Minor, Caucasus species key); Fernández & Ortiz-Sepúlveda, 2019: 732 (Colombia); Mackay & MacKay, 2019: 744 (Colombia species key); Salata et al., 2019: 89 (Camponotus kiesenwetteri group key); Mackay, 2019: 24, 48, 137 (New World subgenera key, New World species complexes key, New World Camponotus (Camponotus) species key); Salata et al., 2020: 547 (Turano-Balkan Camponotus samius complex key); Ward & Boudinot, 2021: 52 (male-based genus diagnosis, key); Heterick, 2021: 69 (Western Australia species key); Rasoamanana & Fisher, 2022: 137 (Camponotus (Mayria) revision), 146 (Camponotus (Mayria) diagnosis), 148 (synoptic list of Camponotus (Mayria) species, including species groups), 150 (Camponotus (Mayria) species group definitions), 162 (key to Camponotus (Mayria) species (minor workers)); Rakotonirina & Fisher, 2022: 1 (Malagasy Camponotus (Myrmosaga) revision); 14 (Malagasy Camponotus (Myrmosaga) worker diagnosis), 19 (synoptic list of Malagasy Camponotus (Myrmosaga)), 21 (key to Malagasy Camponotus (Myrmosaga) minor workers); Perfilieva, 2022: 418 (wing diagnosis); Borowiec & Salata, 2022: 57 (diagnosis), 58 (key to Greece subgenera, key to Greece Camponotus (Camponotus) species), 66 (key to Greece Camponotus (Myrmentoma) species), 110 (key to Greece Camponotus (Tanaemyrmex) species).

Extant: 1,087 valid species, 410 valid subspecies

Fossil: 30 valid species

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