Ward, P. S. 1980a. A systematic revision of the Rhytidoponera impressa group (Hymenoptera: Formicidae) in Australia and New Guinea. Australian Journal of Zoology 28:475-498. PDF

Ward, P. S. 1980b. Genetic variation and population differentiation in the Rhytidoponera impressa group, a species complex of ponerine ants (Hymenoptera: Formicidae). Evolution 34:1060-1076. PDF

Ward, P. S.; Taylor, R. W. 1981. Allozyme variation, colony structure and genetic relatedness in the primitive ant Nothomyrmecia macrops Clark (Hymenoptera: Formicidae). Journal of the Australian Entomological Society 20:177-183. PDF

Ward, P. S. 1981a. Ecology and life history of the Rhytidoponera impressa group. I. Habitats, nest sites, and foraging behavior. Psyche (Cambridge) 88:89-108. PDF

Ward, P. S. 1981b. Ecology and life history of the Rhytidoponera impressa group. II. Colony origin, seasonal cycles, and reproduction. Psyche (Cambridge) 88:109-126. PDF

Ward, P. S. 1983a. Genetic relatedness and colony organization in a species complex of ponerine ants. I. Genotypic and phenotypic composition of colonies. Behavioral Ecology and Sociobiology 12:285-299. PDF

Ward, P. S. 1983b. Genetic relatedness and colony organization in a species complex of ponerine ants. II. Patterns of sex ratio investment. Behavioral Ecology and Sociobiology 12:301-307. PDF

Ward, P. S. 1984. A revision of the ant genus Rhytidoponera (Hymenoptera: Formicidae) in New Caledonia. Australian Journal of Zoology 32:131-175. PDF

Ward, P. S. 1985a. Taxonomic congruence and disparity in an insular ant fauna: Rhytidoponera in New Caledonia. Systematic Zoology 34:140-151. PDF

Ward, P. S. 1985b. The Nearctic species of the genus Pseudomyrmex (Hymenoptera: Formicidae). Quaestiones Entomologicae 21:209-246. PDF

Ward, P. S. 1986. Functional queens in the Australian greenhead ant, Rhytidoponera metallica (Hymenoptera: Formicidae). Psyche (Cambridge) 93:1-12. PDF

Ward, P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55(2):1-16. PDF

Ward, P. S. 1988. Mesic elements in the western Nearctic ant fauna: taxonomic and biological notes on Amblyopone, Proceratium, and Smithistruma (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 61:102-124. PDF

Ward, P. S. 1989a. Systematic studies on pseudomyrmecine ants: revision of the Pseudomyrmex oculatus and P. subtilissimus species groups, with taxonomic comments on other species. Quaestiones Entomologicae 25:393-468. PDF

Ward, P. S. 1989b. Genetic and social changes associated with ant speciation. Pp. 123-148 in: Breed, M. D.; Page, R. E. (eds.) 1989. The genetics of social evolution. Boulder: Westview Press, 213 pp.

Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Systematic Entomology 15:449-489. PDF

Ward, P. S. 1991. Phylogenetic analysis of pseudomyrmecine ants associated with domatia-bearing plants. Pp. 335-352 in: Huxley, C. R.; Cutler, D. F. (eds.) 1991. Ant-plant interactions. Oxford: Oxford University Press, xviii + 601 pp. PDF

Baroni Urbani, C.; Bolton, B.; Ward, P. S. 1992. The internal phylogeny of ants (Hymenoptera: Formicidae). Systematic Entomology 17:301-329. PDF

Ward, P. S. 1992. Ants of the genus Pseudomyrmex (Hymenoptera: Formicidae) from Dominican amber, with a synopsis of the extant Antillean species. Psyche (Cambridge) 99:55-85. PDF

Frumhoff, P. C.; Ward, P. S. 1992. Individual-level selection, colony-level selection, and the association between polygyny and worker monomorphism in ants. American Naturalist 139:559-590. PDF

Ward, P. S. 1993. Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae). Journal of Hymenoptera Research 2:117-168. PDF

Notes: Errata in J. Hym. Res. 3:309-310. (1994.10.15)

Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology 19:159-175. PDF

Ward, P. S. 1995. Ants on the run. [Review of: Gotwald, W. H., Jr. 1995. Army ants: the biology of social predation. Ithaca, New York: Cornell University Press, xviii + 302 pp.]. Science (Washington, D. C.) 270:319-320. PDF

Ward, P. S. 1996. A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants. Systematic Entomology 21:253-263. PDF

Ward, P. S.; Bolton, B.; Shattuck, S. O.; Brown, W. L., Jr. 1996. A bibliography of ant systematics. University of California Publications in Entomology 116:1-417.

Olson, D. M.; Ward, P. S. 1996. The ant fauna (Hymenoptera: Formicidae) of Kirindy Forest (tropical dry forest) in western Madagascar. Pp. 161-164 in: Ganzhorn, J. U.; Sorg, J.-P. (eds.) 1996. Ecology and economy of a tropical dry forest in Madagascar. Primate Report 46-1. Göttingen: German Primate Center (DPZ), 382 pp. PDF

Ward, P. S. 1997. Ant soldiers are not modified queens. Nature (London) 385:494-495. PDF

Brady, S. G.; Gadau, J.; Ward, P. S. 1999. Is the ant genus Camponotus paraphyletic? [Abstract.]. Pp. 27-28 in: Publication Committee 1999. 4th International Hymenopterists Conference, 6-11th January 1999, Canberra, Australia. Program and abstracts. Glen Osmond, South Australia: 4th International Hymenopterists Conference, 96 pp.

Gadau, J.; Brady, S. G.; Ward, P. S. 1999. Systematics, distribution and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92:514-522. PDF

Ward, P. S. 1999a. Deceptive similarity in army ants of the genus Neivamyrmex (Hymenoptera: Formicidae): taxonomy, distribution and biology of N. californicus (Mayr) and N. nigrescens (Cresson). Journal of Hymenoptera Research 8:74-97. PDF