Khalife, A.; Keller, R. A.; Billen, J.; Hita Garcia, F.; Economo, E. P.; Peeters, C. 2018. Skeletomuscular adaptations of head and legs of Melissotarsus ants for tunnelling through living wood. Frontiers in Zoology 15 (30):11 pp. 10.1186/s12983-018-0277-6 PDF

Notes: Melissotarsus anatomy, morphological adaptations

Billen, J.; Khalife, A.; Ito, F.; Anh, N.D.; Esteves, F. A. 2021. The basitarsal sulcus gland, a novel exocrine structure in ants. Arthropod Structure & Development 61: Article 101041:1-10. 10.1016/j.asd.2021.101041 PDF

Notes: Basitarsal sulcus gland anatomy

Khalife, A.; Peeters, C.; Economo, E. P. 2022. Minute workers and large soldiers in the subterranean ant Carebara perpusilla: Musculoskeletal consequences of Haller's rule in the thorax. Arthropod Structure and Development 69:101188. 10.1016/j.asd.2022.101188 PDF

Khalife, A.; Sasaki, H.; Ito, F. 2024. Intercastes in the ant Aphaenogaster rugulosa (Formicidae: Myrmicinae): morphological variation and adaptive potential. Insectes Sociaux 71:11 pp. [online early] 10.1007/s00040-024-00987-y PDF

Sasaki, H.; Khalife, A.; Ito, F. 2025. Colony composition, prey preference and behavioral characteristics of Leptanilla kubotai (Hymenoptera: Formicidae: Leptanillinae). Asian Myrmecology 18:e018001, 9 pp. [online early] 10.20362/am.018001 PDF

Khachonpisitsak, S.; Jaitrong, W.; Jeenthong, T.; Wongkunanusorn, C.; Laedprathom, K.; Chaianunporn, T.; Khalife, A.; Aupanun, S.; Ito, F. 2025. Ant-plants as oases: Colonies of Cladomyrma sirindhornae (Hymenoptera, Formicidae) thrive in their host plants Sphenodesme pentandra in both the wet and dry seasons. Journal of Hymenoptera Research 98:559-577. [online early] 10.3897/jhr.98.144518 PDF

Oberski, J. T.; Griebenow, Z. H.; Adams, R. M. M.; Andersen, A.; Andrade-Silva, J.; Barden, P.; Borowiec, M.; Brady, S.; Casadei-Ferreira, A.; Csősz, S.; Dias, A. M.; Dias, R. K. S.; Feitosa, R. M.; Fernandez, F.; Fisher, B. L.; General, D. E. M.; Gomez, K.; Hammel, J. U.; Hawkes, P. G.; Janda, M.; Khalife, A.; Ladino, N.; Lieberman, Z. E.; Lucky, A.; Menchetti, M.; do Prado, L. P.; Prebus, M. M.; Probst, R. S.; Punnath, A.; Richter, A.; Salata, S.; Sánchez-Restrepo, A. F.; Schifani, E.; Schultz, T. R.; Silva, R. R.; Sosa-Calvo, J.; Tocora, M. C.; Ulysséa, M. A.; van de Kamp, T.; Wang, W. Y.; Williams, J. L.; Camacho, G. P.; Boudinot, B. E. 2025. Ant systematics: past, present, and future. Insect Systematics and Diversity 9 (4), 11:ixaf025, 42 pp. 10.1093/isd/ixaf025 PDF