Liu, S.-P.; Richter, A.; Stoessel, A.; Beutel, R. G. 2019. The mesosomal anatomy of Myrmecia nigrocincta workers and evolutionary transformations in Formicidae (Hymenoptera). Arthropod Systematics and Phylogeny 77:1-19. 10.26049/ASP77-1-2019-01 PDF

Richter, A.; Keller, R. A.; Rosumek, F. B.; Economo, E. P.; Hita Garcia, F.; Beutel, R. G. 2019. The cephalic anatomy of workers of the ant species Wasmannia affinis (Formicidae, Hymenoptera, Insecta) and its evolutionary implications. Arthropod Structure & Development 49:26-49. 10.1016/j.asd.2019-02-002 PDF

Notes: Cranial anatomy of Formicidae, with Wasmannia as exemplar.

Beutel, R. G.; Richter, A.; Keller, R. A.; Hita Garcia, F.; Matsumura, Y.; Economo, E. P.; Gorb, S. N. 2020. Distal leg structures of the Aculeata (Hymenoptera): A comparative evolutionary study of Sceliphron (Sphecidae) and Formica (Formicidae). Journal of Morphology 2020:1-17. [online early] 10.1002/jmor.21133 PDF

Richter, A.; Hita Garcia, F.; Keller, R. A.; Billen, J.; Economo, E. P.; Beutel, R. G. 2020. Comparative analysis of worker head anatomy of Formica and Brachyponera (Hymenoptera: Formicidae). Arthropod Systematics and Phylogeny 78:133-170. 10.26049/ASP78-1-2020-06 PDF

Richter, A.; Hita Garcia, F.; Keller, R. A.; Billen, J.; Katzke, J.; Boudinot, B. E.; Economo, E. P.; Beutel, R. G. 2021. The head anatomy of Protanilla lini (Hymenoptera: Formicidae: Leptanillinae), with a hypothesis of their mandibular movement. Myrmecological News 31:85-114. 10.25849/myrmecol.news_031:85 PDF

Notes: Head anatomy, Protanilla

Boudinot, B. E.; Moosdorf, O. T. D.; Beutel, R. G.; Richter, A. 2021. Anatomy and evolution of the head of Dorylus helvolus (Formicidae: Dorylinae): Patterns of sex- and caste-limited traits in the sausagefly and the driver ant. Journal of Morphology 282:1616-1658. 10.1002/jmor.21410 PDF

Boudinot, B. E.; Richter, A.; Katzke, J.; Chaul, J. C. M.; Keller, R. A.; Economo, E. P.; Beutel, R. G.; Yamamoto, S. 2022. Evidence for the evolution of eusociality in stem ants and a systematic revision of †Gerontoformica (Hymenoptera: Formicidae). Zoological Journal of the Linnean Society 195:1355-1389. 10.1093/zoolinnean/zlab097/6523228 PDF

Aibekova, L.; Boudinot, B. E.; Beutel, R. G.; Richter, A.; Keller, R. A.; Hita Garcia, F.; Economo, E. P. 2022. The skeletomuscular system of the mesosoma of Formica rufa workers (Hymenoptera: Formicidae). Insect Systematics and Diversity 6 (2): 2:1-26. 10.1093/isd/ixac002 PDF

Richter, A.; Boudinot, B.; Yamamoto, S.; Katzke, J.; Beutel, R. G. 2022. The first reconstruction of the head anatomy of a Cretaceous insect, †Gerontoformica gracilis (Hymenoptera: Formicidae), and the early evolution of ants. Insect Systematics and Diversity 6(5), 4:1-80. 10.1093/isd/ixac013 PDF

Boudinot, B. E.; Richter, A. K.; Hammel, J. U.; Szwedo, J.; Bojarski, B.; Perrichot, V. 2022. Genomic-phenomic reciprocal illumination: Desyopone hereon gen. et sp. nov., an exceptional aneuretine-like fossil ant from Ethiopian amber (Hymenoptera: Formicidae: Ponerinae). Insects 13, 796:19 pp. 10.3390/insects13090796 PDF

Richter, A.; Boudinot, B. E.; Hita Garcia, F.; Billen, J.; Economo, E. P.; Beutel, R. G. 2023. Wonderfully weird: the head anatomy of the armadillo ant, Tatuidris tatusia (Hymenoptera: Formicidae: Agroecomyrmecinae), with evolutionary implications. Myrmecological News 33:35-75. 10.25849/myrmecol.news_033:035 PDF

Griebenow, Z. H.; Richter, A.; van de Kamp, T.; Economo, E. P.; Lieberman, Z. E. 2023. Comparative morphology of male genital skeletomusculature in the Leptanillinae (Hymenoptera: Formicidae), with a standardized muscular terminology for the male genitalia of Hymenoptera. Arthropod Systematics and Phylogeny 81:945-1018. 10.3897/asp.81.e104810 PDF

Boudinot, B. E.; Bock, B. L.; Weingardt, M.; Tröger, D.; Batelka, J.; Li, D.; Richter, A.; Pohl, H.; Moosdorf, O. T. D.; Jandausch, K.; Hammel, J. U.; Beutel, R. G. 2024. Et latet et lucet: Discoveries from the Phyletisches Museum amber and copal collection in Jena, Germany. Deutsche Entomologische Zeitschrift 71 (1):111-176. [online early] 10.3897/dez.71.112433 PDF

Griebenow, Z. H.; Richter, A.; Economo, E. P.; Dang, A. V.; Yamada, A. 2025. Four new species of Leptanillinae (Hymenoptera: Formicidae) from northern Vietnam described with phylogenomics and micro-computed tomography. European Journal of Taxonomy 987:98-145. 10.5852/ejt.2025.987.2867 PDF

Fiorentino, G.; Probst, R.; Richter, A.; Economo, E. P.; Barden, P. 2025. A fossil-informed pattern of body size increase and local extinction in Basiceros dirt ants (Hymenoptera: Formicidae). Proceedings of the Royal Society B: Biological Sciences 292 (2045):20242171 (11 pp.). 10.1098/rspb.2024.2171 PDF

Li, D.; Richter, A.; Olivos-Cisneros, L.; van de Kamp, T.; Kronauer, D. J. C.; Beutel, R. G.; Boudinot, B. E. 2025. The larval morphology of the clonal raider ant, Ooceraea biroi (Forel, 1907) (Dorylinae), with broader implications for the Formicidae and Hymenoptera. Myrmecological News 35:161-188. 10.25849/myrmecol.news_035:161 PDF

Aibekova, L.; Richter, A.; Beutel, R. G.; van de Kamp, T.; Economo, E. P.; Griebenow, Z.; Boudinot, B. E. 2025. The mesosoma of Protanilla (Leptanillinae) and the groundplan of the Formicidae (Hymenoptera). Journal of Morphology 286 (7):e70064, 26 pp. 10.1002/jmor.70064 PDF

Oberski, J. T.; Griebenow, Z. H.; Adams, R. M. M.; Andersen, A.; Andrade-Silva, J.; Barden, P.; Borowiec, M.; Brady, S.; Casadei-Ferreira, A.; Csősz, S.; Dias, A. M.; Dias, R. K. S.; Feitosa, R. M.; Fernandez, F.; Fisher, B. L.; General, D. E. M.; Gomez, K.; Hammel, J. U.; Hawkes, P. G.; Janda, M.; Khalife, A.; Ladino, N.; Lieberman, Z. E.; Lucky, A.; Menchetti, M.; do Prado, L. P.; Prebus, M. M.; Probst, R. S.; Punnath, A.; Richter, A.; Salata, S.; Sánchez-Restrepo, A. F.; Schifani, E.; Schultz, T. R.; Silva, R. R.; Sosa-Calvo, J.; Tocora, M. C.; Ulysséa, M. A.; van de Kamp, T.; Wang, W. Y.; Williams, J. L.; Camacho, G. P.; Boudinot, B. E. 2025. Ant systematics: past, present, and future. Insect Systematics and Diversity 9 (4), 11:ixaf025, 42 pp. 10.1093/isd/ixaf025 PDF

Richter, A.; Keller, R. A.; Hita Garcia, F.; van de Kamp, T.; Economo, E. P. 2025. The evolution of novel mouthpart interlocking mechanisms in ants (Hymenoptera: Formicidae). Zoological Journal of the Linnean Society 205 (1):zlaf113, 26 pp. [online early] 10.1093/zoolinnean/zlaf113 PDF