Ito, F.; Higashi, S. 1990. Temporary social parasitism in the enslaving ant species Formica sanguinea Latreille: an important discovery related to the evolution of dulosis in Formica ants. Journal of Ethology 8:33-35.

Ito, F. 1992 ("1991"). Preliminary report on queenless reproduction in a primitive ponerine ant Amblyopone sp. (reclinata group) in West Java, Indonesia. Psyche (Cambridge) 98:319-322.

Ito, F. 1993. Observation of group recruitment to prey in a primitive ponerine ant, Amblyopone sp. (reclinata group) (Hymenoptera: Formicidae). Insectes Sociaux 40:163-167.

Ito, F. 1994a ("1993"). Queenless reproduction in a primitive ponerine ant Amblyopone bellii (Hymenoptera: Formicidae) in southern India. Journal of the New York Entomological Society 101:574-575.

Ito, F. 1994b. Aggressive interactions among gamergates in queenless ponerine ant Amblyopone sp. (reclinata group). P. 247 in: Lenoir, A.; Arnold, G.; Lepage, M. (eds.) 1994. Les insectes sociaux. 12ème congrès de l'Union Internationale pour l'Étude des Insectes Sociaux. Paris, Sorbonne, 21-27 août 1994. Paris: Université Paris Nord, xxiv + 583 pp.

Ito, F. 1995 ("1994"). Colony composition of two Malaysian ponerine ants, Platythyrea tricuspidata and P. quadridenta: sexual reproduction by workers and production of queens (Hymenoptera: Formicidae). Psyche (Cambridge) 101:209-218.

Ito, F. 1996. Colony characteristics of the Indonesian myrmicine ant Myrmecina sp. (Hymenoptera, Formicidae, Myrmicinae): polygynous reproduction by ergatoid queens. Annals of the Entomological Society of America 89:550-554.

Ito, F. 1997. Colony composition and morphological caste differentiation between ergatoid queens and workers in the ponerine ant genus Leptogenys in the Oriental tropics. Ethology Ecology and Evolution 9:335-343.

Ito, F. 1998. Colony composition and specialized predation on millipedes in the enigmatic ponerine ant genus Probolomyrmex (Hymenoptera, Formicidae). Insectes Sociaux 45:79-83.

Ito, F.; Billen, J. P. J. 1998. Larval hemolymph feeding and oophagy: behavior of queens and workers in the primitive ponerine ant Prionopelta kraepelini (Hymenoptera, Formicidae). Belgian Journal of Zoology 128:201-209.

Peeters, C.; Ito, F.; Gobin, B.; Fannes, W.; Billen, J. 1998. Queen-worker divergence in the spermathecae of ponerine and myrmeciine ants. [Abstract.]. P. 369 in: Schwarz, M. P.; Hogendoorn, K. (eds.) 1998. Social insects at the turn of the millennium. Proceedings of the XIII International Congress of IUSSI. Adelaide, Australia. 29 December 1998 - 3 January 1999. Adelaide: XIII Congress of IUSSI, 535 pp.

Terayama, M.; Ito, F.; Gobin, B. 1998. Three new species of the genus Acanthomyrmex Emery (Hymenoptera: Formicidae) from Indonesia, with notes on the reproductive caste and colony composition. Entomological Science 1:257-264. PDF

Wenseleers, T.; Ito, F.; Van Borm, S.; Huybrechts, R.; Volckaert, F.; Billen, J. 1998. Widespread occurrence of the micro-organism Wolbachia in ants. Proceedings of the Royal Society of London. Series B, Biological Sciences 265:1447-1452. PDF

Billen, J.; Ito, F.; Maile, R.; Morgan, E. D. 1998. The mandibular gland, probably the source of the alarm substance in Leptanilla sp. (Hymenoptera, Formicidae). Naturwissenschaften 85:596-597.

Schoeters, E.; Ito, F.; Miyata, H.; Billen, J. P. J. 1999. Aberrant venom glands in Amblyoponini (Formicidae, Ponerinae): morphology, ultrastructure and histochemistry. Acta Zoologica (Stockholm) 80:3-9.

Billen, J.; Gobin, B.; Ito, F. 1999. Fine structure of the postpygidial gland in Aenictus army ants. Acta Zoologica (Stockholm) 80:307-310.

Billen, J.; Ito, F.; Tsuji, K.; Schoeters, E.; Maile, R.; Morgan, E. D. 2000. Structure and chemistry of the Dufour gland in Pristomyrmex ants (Hymenoptera, Formicidae). Acta Zoologica (Stockholm) 81:159-166.

Peeters, C.; Ito, F. 2001. Colony dispersal and the evolution of queen morphology in social Hymenoptera. Annual Review of Entomology 46:601-630.

Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404. PDF

Yamane, S.; Ito, F. 2001. Descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae) from Java. Pp. 390-394 in: Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404. PDF

Tijskens, M.; Ito, F.; Billen, J. P. J. 2002. Novel exocrine glands in the legs of the ponerine ant Amblyopone reclinata (Hymenoptera, Formicidae). Netherlands Journal of Zoology 52:69-75. PDF

Gobin, B.; Ito, F.; Billen, J. P. J. 2003. The subepithelial gland in ants: a novel exocrine gland closely associated with the cuticle surface. Acta Zoologica (Copenhagen) 84:285-291. PDF

Morgan, E. D.; Jungnickel, H.; Keegans, S. J.; Do Nascimento, R. R.; Billen, J.; Gobin, B.; Ito, F. 2003. Comparative survey of abdominal gland secretions of the ant subfamily Ponerinae. Journal of Chemical Ecology 29:95-114. PDF

Billen, J.; Thijs, B.; Ito, F.; Gobin, B. 2005. The pretarsal footprint gland of the ant Amblyopone reclinata (Hymenoptera, Formicidae) and its role in nestmate recruitment. Arthropod Structure and Development 34:111-115. PDF

Gobin, B.; Ito, F.; Peeters, C.; Billen, J. P. J. 2006. Queen-worker differences in spermatheca reservoir of phylogenetically basal ants. Cell and Tissue Research 326:169-178. PDF

Notes: Published online: 14 June 2006

Morgan, E. D.; Jungnickel, H.; Billen, J.; Ito, F.; Bergmann, J.; Gobin, B. 2008. Contents of the exocrine glands of the ant subfamily Cerapachyinae. Biochemical Systematics and Ecology 36:260-265.

Gobin, B.; Ito, F.; Billen, J. P. J. 2008. Degeneration of sperm reservoir and the loss of mating ability in worker ants. Naturwissenschaften 95:1041-1048. PDF

Notes: Published online: 13 August 2008

Ito, F. 2010. Notes on the biology of the Oriental amblyoponine ant Myopopone castanea: Queen-worker dimorphism, worker polymorphism and larval hemolymph feeding by workers (Hymenoptera: Formicidae). Entomological Science 13:199-204. PDF

Ito, F.; Touyama, Y.; Gotoh, A.; Kitahiro, S.; Billen, J. 2010. Thelytokous parthenogenesis by queens in the dacetine ant Pyramica membranifera (Hymenoptera: Formicidae). Naturwissenschaften 97:725-728. PDF

Notes: Published online: 16 June 2010

Billen, J.; Stroobants, Z.; Hashim, R.; Ito, F. 2013. Diversity and morphology of abdominal glands in workers of the ant genus Myopias (Formicidae, Ponerinae). Arthropod Structure and Development 42:165-172. PDF