References by Johnson, R. A.
Longhurst, C.; Johnson, R. A.; Wood, T. G. 1979. Foraging, recruitment and predation by Decamorium uelense (Santschi) (Formicidae: Myrmicinae) on termites in southern Guinea Savannah, Nigeria. Oecologia (Berlin) 38:83-91.
Johnson, R. A. 1995 ("1994"). Distribution and natural history of the workerless inquiline ant Pogonomyrmex anergismus Cole (Hymenoptera: Formicidae). Psyche (Cambridge) 101:257-262.
Johnson, R. A.; Parker, J. D.; Rissing, S. W. 1996. Rediscovery of the workerless inquiline ant Pogonomyrmex colei and additional notes on natural history (Hymenoptera: Formicidae). Insectes Sociaux 43:69-76.
Parker, J. D.; Dowling, T. E.; Johnson, R. A.; Rissing, S. W. 1998. Evolutionary origins of the ant social parasites of the North American seed harvester ants Pogonomyrmex. [Abstract.]. P. 361 in: Schwarz, M. P.; Hogendoorn, K. (eds.) 1998. Social insects at the turn of the millennium. Proceedings of the XIII International Congress of IUSSI. Adelaide, Australia. 29 December 1998 - 3 January 1999. Adelaide: XIII Congress of IUSSI, 535 pp.
Johnson, R. A. 2000a. Seed-harvester ants (Hymenoptera: Formicidae) of North America: an overview of ecology and biogeography. Sociobiology 36:89-122 + 83-88. PDF
Johnson, R. A. 2000b. Seed harvesting species of Pheidole (Hymenoptera: Formicidae). Sociobiology 36:597.
Johnson, R. A. 2001. Biogeography and community structure of North American seed-harvesting ants. Annual Review of Entomology 46:1-29. PDF
Julian, G. E.; Fewell, J. H.; Gadau, J.; Johnson, R. A.; Larrabee, D. 2002. Genetic determination of the queen caste in an ant hybrid zone. Proceedings of the National Academy of Sciences of the United States of America 99:8157-8160.
Helms Cahan, S.; Parker, J. D.; Rissing, S. W.; Johnson, R. A.; Polony, T. S.; Weiser, M. D.; Smith, D. R. 2002. Extreme genetic differences between queens and workers in hybridizing Pogonomyrmex harvester ants. Proceedings of the Royal Society of London. Series B, Biological Sciences 269:1871-1877. PDF
Johnson, R. A.; Ward, P. S. 2002. Biogeography and endemism of ants (Hymenoptera: Formicidae) in Baja California, Mexico: a first overview. Journal of Biogeography 29:1009-1026. PDF
Anderson, K. E.; Gadau, J.; Mott, B. M.; Johnson, R. A.; Altamirano, A.; Strehl, C.; Fewell, J. H. 2006. Distribution and evolution of genetic caste determination in Pogonomyrmex seed-harvester ants. Ecology 87:2171-2184. PDF
Johnson, R. A.; Holbrook, C. T.; Strehl, C.; Gadau, J. 2007. Population and colony structure and morphometrics in the queen dimorphic harvester ant, Pogonomyrmex pima. . Insectes Sociaux 54:77-86. PDF
Rabeling, C.; Cover, S. P.; Johnson, R. A.; Mueller, U. G. 2007. A review of the North American species of the fungus-gardening ant genus Trachymyrmex (Hymenoptera: Formicidae). Zootaxa 1664:1-53. PDF
Johnson, R. A. 2009. [Untitled. Pogonomyrmex mohavensis Johnson, n sp.]. Pp. 308-310 in: Johnson, R. A.; Overson, R. P. 2009. A new North American species of Pogonomyrmex (Hymenoptera: Formicidae) from the Mohave desert of eastern California and western Nevada. Journal of Hymenoptera Research 18:305-314. PDF
Johnson, R. A.; Overson, R. P. 2009. A new North American species of Pogonomyrmex (Hymenoptera: Formicidae) from the Mohave desert of eastern California and western Nevada. Journal of Hymenoptera Research 18:305-314. PDF
Johnson, R. A.; Overson, R. P.; Moreau, C. S. 2013. A new species of seed-harvester ant, Pogonomyrmex hoelldobleri (Hymenoptera: Formicidae), from the Mohave and Sonoran Deserts of North America. Zootaxa 3646:201-227. PDF
Plowes, N. J. R.; Johnson, R. A.; Hölldobler, B. 2013. Foraging behavior in the ant genus Messor (Hymenoptera: Formicidae: Myrmicinae). Myrmecological News 18:33-49. PDF
Johnson, R. A.; Cover, S. P. 2015. A taxonomic revision of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae) on Hispaniola. Zootaxa 3972 (2):231-249. PDF
Johnson, R. A. 2015. A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I. Zootaxa 4029 (1):1-142. 10.11646/zootaxa.4029.1.1 PDF
Johnson, R. A.; Moreau, C. S. 2016. A new ant genus from southern Argentina and southern Chile, Patagonomyrmex (Hymenoptera: Formicidae). Zootaxa 4139:1-31. 10.11646/zootaxa.4139.1.1 PDF
Gray, K. W.; Cover, S. P.; Johnson, R. A.; Rabeling, C. 2018. The dacetine ant Strumigenys arizonica, an apparent obligate commensal of the fungus-growing ant Trachymyrmex arizonensis in southwestern North America. Insectes Sociaux 65:401-410. 10.1007/s00040-018-0625-8 PDF
van Elst, T.; Eriksson, T. H.; Gadau, J.; Johnson, R. A.; Rabeling, C.; Taylor, J. E.; Borowiec, M. L. 2021. Comprehensive phylogeny of Myrmecocystus honey ants highlights cryptic diversity and infers evolution during aridification of the American Southwest. Molecular Phylogenetics and Evolution 155: Article 107036:1-12. 10.1016/j.ympev.2020.107036 PDF

Notes: Phylogeny, divergence dating, ancestral state reconstruction

Johnson, R. A. 2021. A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part II. Zootaxa 5033 (1):1-230. 10.11646/zootaxa.5033.1.1 PDF
Johnson, R. A.; Borowiec, M. L.; Snelling, R. R.; Cole, A. C. 2022. A taxonomic revision and a review of the biology of the North American seed-harvester ant genus Veromessor (Hymenoptera: Formicidae: Myrmicinae). Zootaxa 5206 (1):1-115. 10.11646/zootaxa.5206.1.1 PDF
Johnson, R. A.; Overson, R. P.; Wilson, E. O. 2024. E.O. Wilson’s last quest: are there ants on the Falkland Islands? Dugesiana 31 (2):125-128. PDF
Johnson, R. A. 2025. A new species of seed-harvester ant in the genus Pogonomyrmex (P. brevibarbis group) with ergatoid queens and intercastes from the Andes of Argentina (Hymenoptera: Formicidae: Myrmicinae). Zootaxa 5693 (2):223-242. 10.11646/zootaxa.5693.2.5 PDF
Borowiec, M. L.; Eskew, S. L.; Noble-Stuen, A.; Williams, J. L.; Johnson, R. A. 2025. Evolution of the North American seed-harvester ant genus Veromessor (Hymenoptera: Formicidae). Systematic Entomology -:e70002, 15 pp. [online early] 10.1111/syen.70002 PDF
Protonyms originally described by author
Protonym Taxon Authorship Rank Status
Pogonomyrmex mohavensis Pogonomyrmex mohavensis Johnson, 2009 Species Valid Original combination
Pogonomyrmex hoelldobleri Pogonomyrmex hoelldobleri Johnson et al., 2013 Species Valid Original combination
Pogonomyrmex tinogasta Pogonomyrmex tinogasta Johnson, 2015 Species Valid Original combination
Patagonomyrmex Patagonomyrmex Johnson & Moreau, 2016 Genus Valid
Pogonomyrmex cusquena Pogonomyrmex cusquena Johnson, 2021 Species Valid Original combination
Pogonomyrmex excelsior Pogonomyrmex excelsior Johnson, 2021 Species Valid Original combination
Pogonomyrmex forelii Pogonomyrmex forelii Johnson, 2021 Species Valid Original combination
Pogonomyrmex granulatus Pogonomyrmex granulatus Johnson, 2021 Species Valid Original combination
Pogonomyrmex lagunabravensis Pogonomyrmex lagunabravensis Johnson, 2021 Species Valid Original combination
Pogonomyrmex loaensis Pogonomyrmex loaensis Johnson, 2021 Species Valid Original combination
Pogonomyrmex mapuche Pogonomyrmex mapuche Johnson, 2021 Species Valid Original combination
Pogonomyrmex maulensis Pogonomyrmex maulensis Johnson, 2021 Species Valid Original combination
Pogonomyrmex pichachen Pogonomyrmex pichachen Johnson, 2021 Species Valid Original combination
Pogonomyrmex propinqua Pogonomyrmex propinquus Johnson, 2021 Species Valid Original combination
Pogonomyrmex santschii Pogonomyrmex santschii Johnson, 2021 Species Valid Original combination
Pogonomyrmex strioligaster Pogonomyrmex strioligaster Johnson, 2021 Species Valid Original combination
Pogonomyrmex tafi Pogonomyrmex tafi Johnson, 2021 Species Valid Original combination
Pogonomyrmex varicolor Pogonomyrmex varicolor Johnson, 2021 Species Valid Original combination
Pogonomyrmex wilsoni Pogonomyrmex wilsoni Johnson, 2021 Species Valid Original combination
Pogonomyrmex apterogenos Pogonomyrmex apterogenos Johnson, 2021 Species Valid Original combination
Pogonomyrmex araucania Pogonomyrmex araucania Johnson, 2021 Species Valid Original combination
Pogonomyrmex atacama Pogonomyrmex atacama Johnson, 2021 Species Valid Original combination
Pogonomyrmex bolivianus Pogonomyrmex bolivianus Johnson, 2021 Species Valid Original combination
Pogonomyrmex colca Pogonomyrmex colca Johnson, 2021 Species Valid Original combination
Veromessor pseudolariversi Veromessor pseudolariversi Johnson et al., 2022 Species Valid Original combination
Pogonomyrmex eremicus Pogonomyrmex eremicus Johnson, 2025 Species Valid Original combination
Statistics
Published between 1979–2025
Taxon descriptions between 2009–2025
No. of references 27
No. of described taxa (including combinations) 27
No. of described taxa (valid, including combinations) 26
No. of described taxa (protonyms only) 26
Names belonging to author
Author name No. of references
Johnson, R. A. 27