Lucky, A.; Ward, P. S. 2010. Taxonomic revision of the ant genus Leptomyrmex Mayr (Hymenoptera: Formicidae). Zootaxa 2688:1-67. PDF

Ward, P. S.; Brady, S. G.; Fisher, B. L.; Schultz, T. R. 2010. Phylogeny and biogeography of dolichoderine ants: effects of data partitioning and relict taxa on historical inference. Systematic Biology 59:342-362. PDF

Ward, P. S. 2011. Integrating molecular phylogenetic results into ant taxonomy (Hymenoptera: Formicidae). Myrmecological News 15:21-29. PDF

Ward, P. S.; Sumnicht, T. P. 2012. Molecular and morphological evidence for three sympatric species of Leptanilla (Hymenoptera: Formicidae) on the Greek island of Rhodes. Myrmecological News 17:5-11. PDF

Johnson, B. R.; Borowiec, M. L.; Chiu, J. C.; Lee, E. K.; Atallah, J.; Ward, P. S. 2013. Phylogenomics resolves evolutionary relationships among ants, bees, and wasps. Current Biology 23:2058-2062. PDF

Brady, S. G.; Fisher, B. L.; Schultz, T. R.; Ward, P. S. 2014. The rise of army ants and their relatives: diversification of specialized predatory doryline ants. BMC Evolutionary Biology 14 (93):1-14. 10.1186/1471-2148-14-93 PDF

Ward, P. S. 2014. The phylogeny and evolution of ants. Annual Review of Ecology, Evolution, and Systematics 45:23-43. PDF

Ward, P. S.; Brady, S. G.; Fisher, B. L.; Schultz, T. R. 2015. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology 40:61-81. 10.1111/syen.12090 PDF

Blaimer, B. B.; Brady, S. G.; Schultz, T. R.; Lloyd, M. W.; Fisher, B. L.; Ward, P. S. 2015. Phylogenomic methods outperform traditional multi-locus approaches in resolving deep evolutionary history: a case study of formicine ants. BMC Evolutionary Biology 15 (271):1-14. 10.1186/s12862-015-0552-5 PDF

Chomicki, G.; Ward, P. S.; Renner, S. S. 2015. Macroevolutionary assembly of ant/plant symbioses: Pseudomyrmex ants and their ant-housing plants in the Neotropics. Proceedings of the Royal Society. Series B. 282 (1819):20152200, 55 pp. 10.1098/rspb.2015.2200 PDF

Ward, P. S.; Blaimer, B. B.; Fisher, B. L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex. Zootaxa 4072:343-357. PDF

Boudinot, B. E.; Probst, R. S.; Brandão, C. R. F.; Feitosa, R. M.; Ward, P. S. 2016. Out of the Neotropics: newly discovered relictual species sheds light on the biogeographical history of spider ants (Leptomyrmex, Dolichoderinae, Formicidae). Systematic Entomology 41:658-671. 10.1111/syen.12181 PDF

Ward, P. S.; Fisher, B. L. 2016. Tales of dracula ants: the evolutionary history of the ant subfamily Amblyoponinae (Hymenoptera: Formicidae). Systematic Entomology 41:683-693. 10.1111/syen.12186 PDF

Ward, P. S.; Brady, S. G.; Fisher, B. L.; Schultz, T. R. 2016. Phylogenetic classifications are informative, stable, and pragmatic: the case for monophyletic taxa. Insectes Sociaux 63:489-492. 10.1007/s00040-016-0516-9 PDF

Ward, P. S. 2017. A review of the Pseudomyrmex ferrugineus and Pseudomyrmex goeldii species groups: acacia-ants and relatives (Hymenoptera: Formicidae). Zootaxa 4227 (4):524-542. 10.11646/zootaxa.4227.4.3 PDF

Ward, P. S.; Branstetter, M. G. 2017. The acacia ants revisited: convergent evolution and biogeographic context in an iconic ant/plant mutualism. Proceedings of the Royal Entomological Society of London. Series B 284:1-9. http://dx.doi.org/10.1098/rspb.2016.2569 PDF

Notes: Phylogeny of Pseudomyrmex ferrugineus group

Branstetter, M. G.; Longino, J. T.; Ward, P. S.; Faircloth, B. C. 2017. Enriching the ant tree of life: enhanced UCE bait set for genome-scale phylogenetics of ants and other Hymenoptera. Methods in Ecology and Evolution 8:768-776. 10.1111/2041-210X.12742 PDF

Branstetter, M. G.; Danforth, B. N.; Pitts, J. P.; Faircloth, B. C.; Ward, P. S.; Buffington, M. L.; Gates, M. W.; Kula, R. R.; Brady, S. G. 2017. Phylogenomic insights into the evolution of stinging wasps and the origins of ants and bees. Current Biology 27:1019-1025. 10.1016/j.cub.2017.03.027 PDF

Blaimer, B. B.; Ward, P. S.; Schultz, T. R.; Fisher, B. L.; Brady, S. G. 2018. Paleotropical diversification dominates the evolution of the hyperdiverse ant tribe Crematogastrini (Hymenoptera: Formicidae). Insect Systematics and Diversity 2 (5):1-14. 10.1093/isd/ixy013 PDF

Notes: Phylogeny of Crematogastrini

Ward, P. S. 2019. Subfamilia Pseudomyrmecinae. Capítulo 33. Pp. 1089-1113 in: Fernández, F.; Guerrero, R. J.; Delsinne, T. (eds.) 2019d. Hormigas de Colombia. Bogotá: Universidad Nacional de Colombia, 1198 pp. PDF

Borowiec, M. L.; Rabeling, C.; Brady, S. G.; Fisher, B. L.; Schultz, T. R.; Ward, P. S. 2019. Compositional heterogeneity and outgroup choice influence the internal phylogeny of the ants. Molecular Phylogenetics and Evolution 134:111-121. 10.1016/j.ympev.2019.01.024 PDF

Ward, P. S.; Boudinot, B. E. 2021. Grappling with homoplasy: taxonomic refinements and reassignments in the ant genera Camponotus and Colobopsis (Hymenoptera: Formicidae). Arthropod Systematics and Phylogeny 79:37-56. 10.3897/asp.79.e66978 PDF

Ward, P. S.; Branstetter, M. G. 2022. Species paraphyly and social parasitism: phylogenomics, morphology, and geography clarify the evolution of the Pseudomyrmex elongatulus group (Hymenoptera: Formicidae), a Mesoamerican ant clade. Insect Systematics and Diversity 6 (1): 4:1-31. 10.1093/isd/iaxb025 PDF

Ward, P. S. 2022. The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species. Zootaxa 5102 (1):1-70. 10.11646/zootaxa.5102.1.1 PDF

Ward, P. S.; Blaimer, B. B. 2022. Taxonomy in the phylogenomic era: species boundaries and phylogenetic relationships among North American ants of the Crematogaster scutellaris group (Formicidae: Hymenoptera). Zoological Journal of the Linnean Society 194:893-937. PDF

Romiguier, J.; Borowiec, M. L.; Weyna, A.; Helleu, Q.; Loire, E.; La Mendola, C.; Rabeling, C.; Fisher, B. L.; Ward, P. S.; Keller, L. 2022. Ant phylogenomics reveals a natural selection hotspot preceding the origin of complex eusociality. Current Biology 32:2942-2947. 10.1016/j.cub.2022.05.001 PDF

Mera-Rodríguez, D.; Jourdan, H.; Ward, P. S.; Shattuck, S.; Cover, S. P.; Wilson, E. O.; Rabeling, C. 2023. Biogeography and evolution of social parasitism in Australian Myrmecia bulldog ants revealed by phylogenomics. Molecular Phylogenetics and Evolution 186 (107825):1-18. [online early] 10.1016/j.ympev.2023.107825 PDF

Ward, P. S.; Fisher, B. L.; Wernegreen, J. J.; Blaimer, B. B. 2025. Evolutionary history, novel lineages and symbiont coevolution in the ant tribe Camponotini (Hymenoptera: Formicidae). Systematic Entomology 50 (3):646-676. 10.1111/syen.12678 PDF

Notes: Published "online early" 20 May 2025 with different pagination.