Jackson, B. D.; Keegans, S. J.; Morgan, E. D.; Clark, W. H.; Blom, P. E. 1991. Chemotaxonomic study of undescribed species of Myrmica ant from Idaho. Journal of Chemical Ecology 17:335-342.

Keegans, S. J.; Morgan, E. D.; Agosti, D.; Wehner, R. 1992. What do glands tell us about species? A chemical case study of Cataglyphis ants. Biochemical Systematics and Ecology 20:559-572. PDF

Morgan, E. D. 1992. The chemical diversity of ants: is there system in diversity? Pp. 183-194 in: Billen, J. P. J. (ed.) 1992. Biology and evolution of social insects. Leuven: Leuven University Press, ix + 390 pp.

Morgan, E. D.; Hölldobler, B.; Vaisar, T.; Jackson, B. D. 1992. Contents of poison apparatus and their relation to trail-following in the ant Daceton armigerum. Journal of Chemical Ecology 18:2161-2168.

Keegans, S. J.; Billen, J. P. J.; Morgan, E. D.; Gökcen, O. A. 1993. Volatile glandular secretions of three species of New World army ants, Eciton burchelli, Labidus coecus and Labidus praedator. Journal of Chemical Ecology 19:2705-2719.

Nascimento, R. R. do; Billen, J. P. J.; Morgan, E. D. 1993. The exocrine secretions of the jumping ant Harpegnathos saltator. Comparative Biochemistry and Physiology. B. Comparative Biochemistry 104:505-508. PDF

Nascimento, R. R. do; Jackson, B. D.; Morgan, E. D.; Clark, W. H.; Blom, P. E. 1993. Chemical secretions of two sympatric harvester ants, Pogonomyrmex salinus and Messor lobognathus. Journal of Chemical Ecology 19:1993-2005.

Oldham, N. J.; Morgan, E. D.; Gobin, B.; Billen, J. P. J. 1994. First identification of a trail pheromone of an army ant (Aenictus species). Experientia (Basel) 50:763-765. PDF

Oldham, N. J.; Morgan, E. D.; Gobin, B.; Schoeters, E.; Billen, J. P. J. 1994. Volatile secretions of Old World army ant Aenictus rotundatus and chemotaxonomic implications of army ant Dufour gland chemistry. Journal of Chemical Ecology 20:3297-3305.

Oldham, N. J.; Morgan, E. D.; Hölldobler, B. 1994. The recruitment pheromones of Aphaenogaster (=Novomessor) cockerelli and A. (=Novomessor) albisetosus, two closely related myrmicine ants. P. 486 in: Lenoir, A.; Arnold, G.; Lepage, M. (eds.) 1994. Les insectes sociaux. 12ème congrès de l'Union Internationale pour l'Étude des Insectes Sociaux. Paris, Sorbonne, 21-27 août 1994. Paris: Université Paris Nord, xxiv + 583 pp.

Hölldobler, B.; Oldham, N. J.; Morgan, E. D.; König, W. A. 1995. Recruitment pheromones in the ants Aphaenogaster albisetosus and A. cockerelli (Hymenoptera: Formicidae). Journal of Insect Physiology 41:739-744.

Nascimento, R. R. do; Schoeters, E.; Morgan, E. D.; Billen, J.; Stradling, D. J. 1996. Chemistry of metapleural gland secretions of three attine ants, Atta sexdens rubropilosa, Atta cephalotes, and Acromyrmex octospinosus (Hymenoptera: Formicidae). Journal of Chemical Ecology 22:987-1000.

Agosti, D.; Austin, C.; Gökcen, O. A.; König, W. A.; Morgan, E. D.; Scott, E. D.; Wehner, R. 1996. (S)-2-methyl-1-hexanol, characteristic mandibular gland substance of ants of the Cataglyphis bicolor group. Chemoecology 7:57-60. PDF

Morgan, E. D.; Billen, J. P. J. 1998. Pheromone communication in social insects: sources and secretions. Pp. 3-33 in: Vander Meer, R. K.; Breed, M. D.; Espelie, K. E.; Winston, M. L. (eds.) 1998. Pheromone communication in social insects. Ants, wasps, bees, and termites. Boulder: Westview Press, x + 368 pp.

Billen, J.; Ito, F.; Maile, R.; Morgan, E. D. 1998. The mandibular gland, probably the source of the alarm substance in Leptanilla sp. (Hymenoptera, Formicidae). Naturwissenschaften 85:596-597.

Billen, J.; Ito, F.; Tsuji, K.; Schoeters, E.; Maile, R.; Morgan, E. D. 2000. Structure and chemistry of the Dufour gland in Pristomyrmex ants (Hymenoptera, Formicidae). Acta Zoologica (Stockholm) 81:159-166.

Gobin, B.; Rüppell, O.; Hartmann, A.; Jungnickel, H.; Morgan, E. D.; Billen, J. 2001. A new type of exocrine gland and its function in mass recruitment in the ant Cylindromyrmex whymperi (Formicidae, Cerapachyinae). Naturwissenschaften 88:395-399.

Morgan, E. D.; Jungnickel, H.; Keegans, S. J.; Do Nascimento, R. R.; Billen, J.; Gobin, B.; Ito, F. 2003. Comparative survey of abdominal gland secretions of the ant subfamily Ponerinae. Journal of Chemical Ecology 29:95-114. PDF

Morgan, E. D.; Jungnickel, H.; Billen, J.; Ito, F.; Bergmann, J.; Gobin, B. 2008. Contents of the exocrine glands of the ant subfamily Cerapachyinae. Biochemical Systematics and Ecology 36:260-265.

Morgan, E. D. 2009. Trail pheromones of ants. Physiological Entomology 34:1-17. PDF