<ref name="Radchenko_&_Proshchalykin_2021">{{cite journal |first1=A. G. |last1=Radchenko |first2=M. Y. |last2=Proshchalykin |year=2021 |title=New extinct ant genus (Hymenoptera, Formicidae, Myrmicinae) from late Eocene Rovno amber |url=https://antcat.org/documents/8147/radchenko___proshchalykin_2021_jhymres_myrmicinae_lelejus_gen_nov_rovno.pdf |journal=Journal of Hymenoptera Research |publisher= |volume=84 |issue= |pages=271–282 |doi=10.3897/jhr.84.65238 }}</ref>