<ref name="Radchenko_et_al_2018">{{cite journal |first1=A. G. |last1=Radchenko |first2=G. M. |last2=Dlussky |first3=K. S. |last3=Perfilieva |year=2018 |title=A new extinct ant genus (Hymenoptera: Formicidae: Myrmicinae) from the Late Eocene Rovno amber – a putative ancestor of the ''Leptothorax'' genus group |url=https://antcat.org/documents/8188/radchenko_etal_2018_ba_proleptothorax.pdf |journal=Myrmecological News |publisher= |volume=27 |issue= |pages=111–117 |doi=10.25849/myrmecol.news_027:111 }}</ref>