<ref name="Boudinot_2015">{{cite journal |first1=B. E. |last1=Boudinot |year=2015 |title=Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): a new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages |url=https://antcat.org/documents/6385/242-article_text-1337-1-10-20150420.pdf |journal=European Journal of Taxonomy |publisher= |volume=120 |issue= |pages=1–62 |doi= }}</ref>