<ref name="Moreau_2011">{{cite journal |first1=C. S. |last1=Moreau |year=2011 |title=What do molecular clocks tell us about the evolution of ants? |url=https://antcat.org/documents/5248/Moreau_2011_Am_Entomol molecular clocks.pdf |journal=American Entomologist |publisher= |volume=57 |issue= |pages=52–53 |doi= }}</ref>