<ref name="Yamane_&_Hashimoto_1999">{{cite journal |first1=S. |last1=Yamane |first2=Y. |last2=Hashimoto |year=1999 |title=A remarkable new species of the army ant genus ''Aenictus'' (Hymenoptera, Formicidae) with a polymorphic worker caste |url= |journal=Tropics |publisher= |volume=8 |issue= |pages=427–432 |doi= }}</ref>