<ref name="Linksvayer_et_al_2006">{{cite journal |first1=T. A. |last1=Linksvayer |first2=M. J. |last2=Wade |first3=D. M. |last3=Gordon |year=2006 |title=Genetic caste determination in harvester ants: possible origin and maintenance by cyto-nuclear epistasis |url=https://antcat.org/documents/4509/Linksvayer_et_al_2006_Ecology.pdf |journal=Ecology |publisher= |volume=87 |issue= |pages=2185–2193 |doi= }}</ref>