<ref name="Lattke_1999">{{cite journal |first1=J. E. |last1=Lattke |year=1999 |title=A new species of fungus-growing ant and its implications for attine phylogeny (Hymenoptera: Formicidae) |url=https://antcat.org/documents/3934/8392.pdf |journal=Systematic Entomology |publisher= |volume=24 |issue= |pages=1–6 |doi= }}</ref>