<ref name="Johnson_et_al_2007">{{cite journal |first1=R. A. |last1=Johnson |first2=C. T. |last2=Holbrook |first3=C. |last3=Strehl |first4=J. |last4=Gadau |year=2007 |title=Population and colony structure and morphometrics in the queen dimorphic harvester ant, ''Pogonomyrmex pima. '' |url=https://antcat.org/documents/4462/Johnson_et_al_2007_Insectes_Soc.pdf |journal=Insectes Sociaux |publisher= |volume=54 |issue= |pages=77–86 |doi= }}</ref>