<ref name="Janda_et_al_2004">{{cite journal |first1=M. |last1=Janda |first2=D. |last2=Folková |first3=J. |last3=Zrzavý |year=2004 |title=Phylogeny of ''Lasius'' ants based on mitochondrial DNA and morphology, and the evolution of social parasitism in the Lasiini (Hymenoptera: Formicidae) |url=https://antcat.org/documents/4453/Janda_et_al_2004_MPE.pdf |journal=Molecular Phylogenetics and Evolution |publisher= |volume=33 |issue= |pages=595–614 |doi= }}</ref>