<ref name="Heinze_et_al_2005">{{cite journal |first1=J. |last1=Heinze |first2=A. |last2=Trindl |first3=B. |last3=Seifert |first4=K. |last4=Yamauchi |year=2005 |title=Evolution of male morphology in the ant genus ''Cardiocondyla'' |url=https://antcat.org/documents/4428/Heinze_et_al_2005_Mol_Phylogenet_Evol.pdf |journal=Molecular Phylogenetics and Evolution |publisher= |volume=37 |issue= |pages=278–288 |doi= }}</ref>