<ref name="Heinze_et_al_2004">{{cite journal |first1=J. |last1=Heinze |first2=A. |last2=Böttcher |first3=S. |last3=Cremer |year=2004 |title=Production of winged and wingless males in the ant, ''Cardiocondyla minutior'' |url=https://antcat.org/documents/4427/Heinze_et_al_2004_Insectes_Soc.pdf |journal=Insectes Sociaux |publisher= |volume=51 |issue= |pages=275–278 |doi= }}</ref>