<ref name="Fowler_et_al_2000">{{cite journal |first1=H. G. |last1=Fowler |first2=J. H. C. |last2=Delabie |first3=P. R. S. |last3=Moutinho |year=2000 |title=Hypogaeic and epigaeic ant (Hymenoptera: Formicidae) assemblages of Atlantic costal rainforest and dry mature and secondary Amazon forest in Brazil: continuums or communities |url=https://antcat.org/documents/8042/fowler_et_al-_2000-_hypogaeic_and_epigaeic_ant_-hymenoptera__formicidae-_assemblages_of_atlantic_costal_rainforest_and_dry_mature_and_secondary_amazon_forest_in_brazil__continuums_or_communities.pdf |journal=Tropical Ecology |publisher= |volume=41 |issue= |pages=73–80 |doi= }}</ref>