<ref name="Dlussky_&_Rasnitsyn_2003">{{cite journal |first1=G. M. |last1=Dlussky |first2=A. P. |last2=Rasnitsyn |year=2003 |title=Ants (Hymenoptera: Formicidae) of Formation Green River and some other Middle Eocene deposits of North America |url=https://antcat.org/documents/4327/Dlussky&Rasnitsyn_2003_Russ_Entomol_J.pdf |journal=Russian Entomological Journal |publisher= |volume=11 |issue= |pages=411–436 |doi= }}</ref>