<ref name="Yamauchi_1979">{{cite journal |first1=K. |last1=Yamauchi |year=1979 |title=Taxonomical and ecological studies on the ant genus ''Lasius'' in Japan (Hymenoptera: Formicidae). I. Taxonomy |url=https://antcat.org/documents/3748/3521.pdf |journal=Science Report of the Faculty of Education, Gifu University (Natural Science) |publisher= |volume=6 |issue= |pages=147–181 |doi= }}</ref>