<ref name="Yamane_et_al_1994">{{cite journal |first1=S. |last1=Yamane |first2=T. |last2=Iwai |first3=H. |last3=Watanabe |first4=Y. |last4=Yamanouchi |year=1994 |title=Ant fauna of the Tokara Islands, northern Ryukyus, Japan (Hymenoptera, Formicidae) |url= |journal=WWF (Worldwide Fund for Nature) Japan Science Report |publisher= |volume=2 |issue=2 |pages=311–327 |doi= }}</ref>