<ref name="Wilson_1976">{{cite journal |first1=E. O. |last1=Wilson |year=1976 |title=Behavioral discretization and number of castes in an ant species |url=https://antcat.org/documents/3705/13028.pdf |journal=Behavioral Ecology and Sociobiology |publisher= |volume=1 |issue= |pages=141–154 |doi= }}</ref>