<ref name="Wheeler_&_Darlington_1930">{{cite journal |first1=W. M. |last1=Wheeler |first2=P. J., Jr. |last2=Darlington |year=1930 |title=Ant-tree notes from Rio Frio, Colombia |url=http://antbase.org/ants/publications/3461/3461.pdf |journal=Psyche (Cambridge) |publisher= |volume=37 |issue= |pages=107–117 |doi= }}</ref>