<ref name="Wheeler_1933">{{cite journal |first1=W. M. |last1=Wheeler |year=1933 |title=''Mermis'' parasitism in some Australian and Mexican ants |url=https://antcat.org/documents/3615/3426.pdf |journal=Psyche (Cambridge) |publisher= |volume=40 |issue= |pages=20–31 |doi= }}</ref>