<ref name="Wheeler_1925">{{cite journal |first1=W. M. |last1=Wheeler |year=1925 |title=Zoological results of the Swedish Expedition to Central Africa 1921. Insecta. 10. Formicidae |url=https://antcat.org/documents/5199/3380.pdf |journal=Arkiv för Zoologi |publisher= |volume=17A |issue=25 |pages=1–3 |doi= }}</ref>