<ref name="Wheeler_&_Wheeler_1980">{{cite journal |first1=G. C. |last1=Wheeler |first2=J. |last2=Wheeler |year=1980 |title=Supplementary studies on ant larvae: Ponerinae, Myrmicinae and Formicinae |url=https://antcat.org/documents/3296/3128.pdf |journal=Transactions of the American Entomological Society |publisher= |volume=106 |issue= |pages=527–545 |doi= }}</ref>