<ref name="Wheeler_&_Wheeler_1956">{{cite journal |first1=G. C. |last1=Wheeler |first2=J. |last2=Wheeler |year=1956 |title=The ant larvae of the subfamily Pseudomyrmecinae (Hymenoptera: Formicidae) |url=https://antcat.org/documents/3293/3093.pdf |journal=Annals of the Entomological Society of America |publisher= |volume=49 |issue= |pages=374–398 |doi= }}</ref>