<ref name="Shyamalanath_&_Forbes_1984">{{cite journal |first1=S. |last1=Shyamalanath |first2=J. |last2=Forbes |year=1984 |title=Anatomy and histology of the male reproductive system in the adult and pupa of the doryline ant, ''Aenictus'' ''gracilis'' Emery (Hymenoptera: Formicidae) |url= |journal=Journal of the New York Entomological Society |publisher= |volume=91 |issue= |pages=377–393 |doi= }}</ref>