<ref name="Seppä_1992">{{cite journal |first1=P. |last1=Seppä |year=1992 |title=Genetic relatedness of worker nestmates in ''Myrmica'' ''ruginodis'' (Hymenoptera: Formicidae) populations |url= |journal=Behavioral Ecology and Sociobiology |publisher= |volume=30 |issue= |pages=253–260 |doi= }}</ref>