<ref name="Rosengren_et_al_1994">{{cite journal |first1=R. |last1=Rosengren |first2=D. |last2=Chautems |first3=D. |last3=Cherix |first4=W. |last4=Fortelius |first5=L. |last5=Keller |year=1994 |title=Separation of two sympatric sibling species of ''Formica'' L. ants by a behavioural choice test based on brood discrimination |url=https://antcat.org/documents/2450/13322.pdf |journal=Memorabilia Zoologica |publisher= |volume=48 |issue= |pages=237–249 |doi= }}</ref>