<ref name="Ogata_et_al_1995">{{cite journal |first1=K. |last1=Ogata |first2=M. |last2=Terayama |first3=K. |last3=Masuko |year=1995 |title=The ant genus ''Leptanilla'': discovery of the worker-associated male of ''L. japonica'', and a description of a new species from Taiwan (Hymenoptera: Formicidae: Leptanillinae) |url=https://antcat.org/documents/2297/8549.pdf |journal=Systematic Entomology |publisher= |volume=20 |issue= |pages=27–34 |doi= }}</ref>