<ref name="MacKay_1993">{{cite journal |first1=W. P. |last1=MacKay |year=1993 |title=The status of the ant ''Leptothorax'' ''pergandei'' Emery (Hymenoptera: Formicidae) |url=https://antcat.org/documents/2072/4405.pdf |journal=Sociobiology |publisher= |volume=21 |issue= |pages=287–297 |doi= }}</ref>