<ref name="Kugler_1984">{{cite journal |first1=J. |last1=Kugler |year=1984 |title=The males of ''Cardiocondyla'' Emery (Hymenoptera: Formicidae) with the description of the winged male of ''Cardiocondyla wroughtoni'' (Forel) |url=https://antcat.org/documents/1873/4658.pdf |journal=Israel Journal of Entomology |publisher= |volume=17 |issue= |pages=1–21 |doi= }}</ref>