<ref name="Imai_et_al_1994">{{cite journal |first1=H. T. |last1=Imai |first2=R. W. |last2=Taylor |first3=R. H. |last3=Crozier |year=1994 |title=Experimental bases for the minimum interaction theory. I. Chromosome evolution in ants of the ''Myrmecia'' ''pilosula'' species complex (Hymenoptera: Formicidae: Myrmeciinae) |url=https://antcat.org/documents/1695/21646.pdf |journal=Japanese Journal of Genetics |publisher= |volume=69 |issue= |pages=137–182 |doi= }}</ref>