<ref name="Hung_et_al_1972">{{cite journal |first1=A. C. F. |last1=Hung |first2=H. T. |last2=Imai |first3=M. |last3=Kubota |year=1972 |title=The chromosomes of nine ant species (Hymenoptera: Formicidae) from Taiwan, Republic of China |url=https://antcat.org/documents/1678/4814.pdf |journal=Annals of the Entomological Society of America |publisher= |volume=65 |issue= |pages=1023–1025 |doi= }}</ref>