<ref name="Hölldobler_et_al_1989">{{cite journal |first1=B. |last1=Hölldobler |first2=J. M. |last2=Palmer |first3=K. |last3=Masuko |first4=W. L., Jr. |last4=Brown |year=1989 |title=New exocrine glands in the legionary ants of the genus ''Leptanilla'' (Hymenoptera: Formicidae: Leptanillinae) |url= |journal=Zoomorphology (Berlin) |publisher= |volume=108 |issue= |pages=255–262 |doi= }}</ref>