<ref name="Hölldobler_&_Lumsden_1980">{{cite journal |first1=B. |last1=Hölldobler |first2=C. J. |last2=Lumsden |year=1980 |title=Territorial strategies in ants |url=https://antcat.org/documents/8270/holldobler_lumsden_1980_science_territorial_strategies_in_ants.pdf |journal=Science (Washington, D. C.) |publisher= |volume=210 |issue= |pages=732–739 |doi= }}</ref>