<ref name="Fortelius_et_al_1987">{{cite journal |first1=W. |last1=Fortelius |first2=P. |last2=Pamilo |first3=R. |last3=Rosengren |first4=L. |last4=Sundström |year=1987 |title=Male size dimorphism and alternative reproductive tactics in ''Formica'' ''exsecta'' ants (Hymenoptera, Formicidae) |url= |journal=Annales Zoologici Fennici |publisher= |volume=24 |issue= |pages=45–54 |doi= }}</ref>