<ref name="Elmes_1991">{{cite journal |first1=G. W. |last1=Elmes |year=1991 |title=Mating strategy and isolation between the two forms, ''macrogyna'' and ''microgyna'', of ''Myrmica ruginodis'' (Hym. Formicidae) |url= |journal=Ecological Entomology |publisher= |volume=16 |issue= |pages=411–423 |doi= }}</ref>