<ref name="Donisthorpe_1922">{{cite journal |first1=H. |last1=Donisthorpe |year=1922 |title=The subfamilies of Formicidae |url=https://antcat.org/documents/920/5757.pdf |journal=Transactions of the Entomological Society of London |publisher= |volume=1921 |issue= |pages=xl–xlvi |doi= }}</ref>