<ref name="Davies_et_al_1994">{{cite journal |first1=S. J. |last1=Davies |first2=M. H. |last2=Villet |first3=T. M. |last3=Blomefield |first4=R. M. |last4=Crewe |year=1994 |title=Reproduction and division of labour in ''Leptogenys schwabi'' Forel (Hymenoptera Formicidae), a polygynous, queenless ponerine ant |url= |journal=Ethology Ecology and Evolution |publisher= |volume=6 |issue= |pages=507–517 |doi= }}</ref>