<ref name="Brener_&_Ruggiero_1994">{{cite journal |first1=A. G. F. |last1=Brener |first2=A. |last2=Ruggiero |year=1994 |title=Leaf-cutting ants (''Atta'' and ''Acromyrmex'') inhabiting Argentina: patterns in species richness and geographic range sizes |url= |journal=Journal of Biogeography |publisher= |volume=21 |issue= |pages=391–399 |doi= }}</ref>