<ref name="Bolton_1977">{{cite journal |first1=B. |last1=Bolton |year=1977 |title=The ant tribe Tetramoriini (Hymenoptera: Formicidae). The genus ''Tetramorium'' Mayr in the Oriental and Indo-Australian regions, and in Australia |url=https://antcat.org/documents/238/6434.pdf |journal=Bulletin of the British Museum (Natural History). Entomology |publisher= |volume=36 |issue= |pages=67–151 |doi= }}</ref>