<ref name="Berndt_&_Kremer_1984">{{cite journal |first1=K.-P. |last1=Berndt |first2=G. |last2=Kremer |year=1984 |title=New categories in the gynandromorphism of ants |url= |journal=Insectes Sociaux |publisher= |volume=30 |issue= |pages=461–465 |doi= }}</ref>