<ref name="Bagnères_et_al_1991">{{cite journal |first1=A.-G. |last1=Bagnères |first2=E. D. |last2=Morgan |first3=J.-L. |last3=Clement |year=1991 |title=Species-specific secretions of the Dufour glands of three species of formicine ants (Hymenoptera, Formicidae) |url= |journal=Biochemical Systematics and Ecology |publisher= |volume=19 |issue= |pages=25–33 |doi= }}</ref>