<ref name="Angus_et_al_1993">{{cite journal |first1=C. J. |last1=Angus |first2=M. K. |last2=Jones |first3=A. J. |last3=Beattie |year=1993 |title=A possible explanation for size differences in the metapleural glands of ants (Hymenoptera: Formicidae) |url= |journal=Journal of the Australian Entomological Society |publisher= |volume=32 |issue= |pages=73–77 |doi= }}</ref>