<ref name="Agosti_&_Bolton_1990">{{cite journal |first1=D. |last1=Agosti |first2=B. |last2=Bolton |year=1990 |title=New characters to differentiate the ant genera ''Lasius'' F. and ''Formica'' L. (Hymenoptera: Formicidae) |url=https://antcat.org/documents/20/6855.pdf |journal=Entomologist's Gazette |publisher= |volume=41 |issue= |pages=149–156 |doi= }}</ref>